Toto the elephant has died aged 46 at Blair Drummond Safari Park.
The gentle giant had to be put to sleep after she collapsed in her pen.
An emotional Facebook message said: "Staff tried to get her back onto her feet but with no success and sadly after the park veterinarians’ thorough examination a difficult decision was made to euthanise Toto to avoid continued stress and suffering.
"Over the years our elephant team have worked tirelessly, improving the welfare standards of our elephants, ensuring they receive the best daily care particularly during the later stages of their lives.
"Her keepers are devastated but they all understand that this was the last option and necessary and most humane course of action to take."
Toto was rescued from the wild and was a firm favourite at the safari park, near Stirling, over the last 19 years.
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