Edinburgh Zoo has said it will attempt to breed its giant pandas again next year after confirming female Tian Tian is no longer pregnant.
Tian Tian's hormones have returned to normal and it is likely that a reabsorption of the foetus occurred, the zoo said.
Keepers described it as "devastating" news and said the breeding programme will continue.
Iain Valentine, director of giant pandas for the Royal Zoological Society of Scotland, said: "First we need to make sure that Tian Tian and Yang Guang are in great condition.
"Tian Tian still needs to get back into a routine, which will take a few months, and then from about the shortest day we will start to monitor her again for breeding."
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